Philadelphia, PA -- (Sports Network) - Appalachian State retained the No. 1 ranking in The Sports Network FCS Top-25 Poll, despite its 41-13 loss to defending BCS champion LSU, but its lead at the top of the poll closed.

The Mountaineers received 62 of 110 first-place votes and 2,625 points after getting 97 first-place votes in the preseason poll.

North Dakota State, a 41-6 winner over Austin Peay, remained second with 42 first-place votes, up from one first-place vote in the preseason poll. The Bison had 2,555 total points.

Other teams receiving first-place votes were No. 3 Richmond with four, No. 5 Montana and No. 11 Cal Poly with one each. James Madison, which had three first-place votes in the preseason poll, didn't have any first-place votes after losing 31-7 to Duke and fell from sixth to eighth in the new poll.

There was plenty of shuffling after the opening week of action. Richmond moved up one spot from fourth to third after its 28-10 victory at then-No. 15 Elon, replacing Northern Iowa, which fell to No. 9 after a 41-17 loss at Brigham Young.

Massachusetts survived an upset bid by Albany to win 28-16 and moved up a slot to No. 4, while Montana moved up from eighth to fifth without playing a game. Montana travels to Cal Poly on Saturday night in a battle between ranked teams.

Delaware's 14-7 loss to Maryland impressed voters enough to move the Blue Hens from ninth to sixth. Eastern Washington's competitive 49-24 loss at Texas Tech didn't hurt the Eagles, who remained seventh. McNeese State fell 35-27 to North Carolina and stayed 10th in the poll.

Cal Poly's 29-27 victory over San Diego State lifted the Mustangs from 14th to 11th. Southern Illinois fell one spot in its bye week from 11th to 12th, Wofford remained 13th after defeating Presbyterian 38-21 and Youngstown State dropped from 12th to 14th after being shut out by Ohio State 43-0.

Elon's loss to Richmond cost it two spots in the poll, the Phoenix falling from 15th to 17th. New Hampshire and Georgia Southern benefited by moving up one slot each, the Wildcats to No. 15 on an off week and Georgia Southern to No. 16 after a 45-21 loss to Georgia.

Western Illinois' near upset of Arkansas caused it to rise from No. 25 to No. 18. The Leathernecks led nearly the entire game before Arkansas scored two touchdowns in the final quarter to win 28-24.

Eastern Illinois slid one position from No. 18 to No. 19 following a 31-12 loss to Central Michigan. The Citadel climbed three places to No. 20 with a 54-7 victory over Webber

The final five positions in the new poll are represented by South Dakota State, Villanova, Central Arkansas, Eastern Kentucky and Harvard.

South Dakota State fell two spots to No. 21 after a 44-17 loss to Iowa State, while Central Arkansas remained at No. 22 by beating Henderson State 38-14.

Villanova, which lost 48-21 to West Virginia, slipped from No. 21 to No. 23 and Eastern Kentucky fell four slots after a 40-7 loss to Cincinnati. Harvard, which doesn't begin play until Sept. 20, also lost a position.